Pierre ex Prain & Burkill, J. Proc. Asiat. Soc. Bengal 10: 13 (1914)
Climber to 10 m. Rhizome c. 4–8 by 1–5 cm, branching and spreading, shallowly buried, periderm hard but lacking rigid, spine-like roots. Indumentum absent. Stems 2–5 mm in diam., twining to the left, annual, unarmed, terete. Leaves simple, alternate, blades ovate to broadly so, membranous to thinly chartaceous and usually translucent when dried, 7–9(–11)-veined, base shallowly to deeply cordate apex acuminate, margins usually deeply 3-lobed, sometimes shallowly so or entire (reproductive shoots only) or with additional lobes (towards stem base); petioles 2–13 cm long; cataphylls and bulbils absent; lateral nodal spines usually present on either side of each node, sometimes absent (especially on distal shoots) or broken off in specimen preparation, curved, hard and rather brittle, to 4 mm long. Inflorescences spicate, pendent, axes sometimes finely tuberculate, tepals inserted on a cup-shaped torus, erect, apices not recurved, fused for 1/2 to 2/3 of their length. Male inflorescences simple or compound, compound inflorescences 1(–2) per axil, simple/partial inflorescences (Pl. 17C) 1–2(–3) per axil, peduncles 2.2–25 mm long, axes 4–33.5 cm long, with an apparently sessile cymule of (1–)2–3 flowers at each node; female inflorescences 1(–2) per axil, simple. Male flowers 0.8–1.4 mm in diam. at anthesis, outer tepals 1.9–2.3 by 0.6–1, narrowly obovate to obovate-oblong, inner tepals 1.9–2.2 by 0.6–0.8 mm, obovate to obovate-oblong, stamens 6. Capsules 18–21 by 25–40 mm. Seeds 3–6 by 5–7 mm, ovoid-lenticular; wings 17–18 by 15–17 mm, extending all around seed margin.